I just looked at your pack and one tip I would suggest is spread out their dominance. Even your alphas, don't max them both out one could 199 the other 200 for example. At fed/watered/ healed your happniness is only about 57 right now. So many wolves have zero dominance that they are 'angry/fighting each other" they don't know where they lie in the pack.
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I spread in increments of 5 or 10 when starting. Just no two wolves can have same number dominance. You can even part them by 1 number. I suggest a small gap incase you want to rearrange wolves. This will incredily boost your overall happiness and your wolves will heal really fast!
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My wolves heal quite fast and my happiness is usually 87
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Side note, besides spreading the dominance. I have discovered that the less den spaces you have available, your pack will get a litte upset when it starts getting down to less then 3 available spaces. I have only noticed this when in the past I'd be out of den space, my pack happiness would drop, even with spread dominance it was inevitable